Grazing (behaviour)
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Grazing is a method of
feeding Eating (also known as consuming) is the ingestion of food, typically to provide a heterotrophic organism with energy and to allow for growth. Animals and other heterotrophs must eat in order to survive — carnivores eat other animals, herbi ...
in which a
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
feeds on low-growing
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
s such as
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
es or other multicellular organisms, such as
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
. Many
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of animals can be said to be grazers, from large animals such as hippopotamuses to small aquatic snails. Grazing
behaviour Behavior (American English) or behaviour (British English) is the range of actions and mannerisms made by individuals, organisms, systems or artificial entities in some environment. These systems can include other systems or organisms as wel ...
is a type of feeding strategy within the
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
of a species. Specific grazing strategies include graminivory (eating grasses);
coprophagy Coprophagia () or coprophagy () is the consumption of feces. The word is derived from the grc, κόπρος , "feces" and , "to eat". Coprophagy refers to many kinds of feces-eating, including eating feces of other species (heterospecifics), of ...
(producing part-digested pellets which are reingested);
pseudoruminant Pseudoruminant is a classification of animals based on their digestive tract differing from the ruminants. Hippopotami and camels are ungulate mammals with a three-chambered stomach (ruminants have a four-chambered stomach) while equids (horses, as ...
(having a multi-chambered stomach but not
chewing the cud Ruminants (suborder Ruminantia) are hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The ...
); and grazing on plants other than grass, such as on
marine algae Marine primary production is the chemical synthesis in the ocean of organic compounds from atmospheric or dissolved carbon dioxide. It principally occurs through the process of photosynthesis, which uses light as its source of energy, but it al ...
. Grazing's ecological effects can include redistributing nutrients, keeping grasslands open or favouring a particular species over another.


Ecology

Many small selective herbivores follow larger grazers which skim off the highest, tough growth of grasses, exposing tender shoots. For terrestrial animals, grazing is normally distinguished from
browsing Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
in that grazing is eating grass or
forb A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
s, whereas browsing is eating woody twigs and
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
from
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s and
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s. Grazing differs from
predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
because the organism being grazed upon may not be killed. It differs from
parasitism Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of lif ...
because the two organisms
live together "Live Together" is a song recorded by British singer, songwriter and actress Lisa Stansfield for her debut album, '' Affection'' (1989). It was written by Stansfield, Ian Devaney and Andy Morris, and produced by Devaney and Morris. It was releas ...
in a constant state of physical externality (i.e. low intimacy).
Water animal Water (chemical formula ) is an inorganic, transparent, tasteless, odorless, and nearly colorless chemical substance, which is the main constituent of Earth's hydrosphere and the fluids of all known living organisms (in which it acts as a ...
s that feed by rasping algae and other micro-organisms from stones are called ''grazers–scrapers''.


Graminivory

Graminivory is a form of grazing involving feeding primarily on grass (specifically "true" grasses in the
Poaceae Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
).
Horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...
s,
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult mal ...
,
capybara The capybaraAlso called capivara (in Brazil), capiguara (in Bolivia), chigüire, chigüiro, or fercho (in Colombia and Venezuela), carpincho (in Argentina, Paraguay and Uruguay) and ronsoco (in Peru). or greater capybara (''Hydrochoerus hydro ...
,
hippopotamus The hippopotamus ( ; : hippopotamuses or hippopotami; ''Hippopotamus amphibius''), also called the hippo, common hippopotamus, or river hippopotamus, is a large semiaquatic mammal native to sub-Saharan Africa. It is one of only two extan ...
es,
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grasshopp ...
s,
geese A goose (plural, : geese) is a bird of any of several waterfowl species in the family (biology), family Anatidae. This group comprises the genera ''Anser (bird), Anser'' (the grey geese and white geese) and ''Branta'' (the black geese). Some o ...
, and
giant panda The giant panda (''Ailuropoda melanoleuca''), also known as the panda bear (or simply the panda), is a bear species endemic to China. It is characterised by its bold black-and-white coat and rotund body. The name "giant panda" is sometimes us ...
s are graminivores. Giant pandas (''Ailuropoda melanoleuca'') are obligate
bamboo Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, bu ...
grazers, 99% of their diet consisting of sub-alpine bamboo species.


Coprophagy

Rabbits are herbivores that feed by grazing on grass,
forb A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
s, and leafy weeds. They graze heavily and rapidly for about the first half-hour of a grazing period (usually in the late afternoon), followed by about half an hour of more selective feeding. If the environment is relatively non-threatening, the rabbit remains outdoors for many hours, grazing at intervals. Their diet contains large amounts of
cellulose Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell wall ...
, which is hard to digest. Rabbits solve this problem by using a form of
hindgut fermentation Hindgut fermentation is a digestive process seen in monogastric herbivores, animals with a simple, single-chambered stomach. Cellulose is digested with the aid of symbiotic bacteria.coprophagy Coprophagia () or coprophagy () is the consumption of feces. The word is derived from the grc, κόπρος , "feces" and , "to eat". Coprophagy refers to many kinds of feces-eating, including eating feces of other species (heterospecifics), of ...
). Rabbits reingest their own droppings (rather than chewing the cud as do cattle and many other grazers) to digest their food further and extract sufficient nutrients. Capybara (''Hydrochoerus hydrochaeris'') are herbivores that graze mainly on grasses and aquatic plants,Capybara Facts.
Smithsonian National Zoological Park. Retrieved on December 16, 2007.
PDF
/ref> as well as fruit and tree Bark (botany), bark.Capybara.
Palm Beach Zoo. Retrieved on December 17, 2007.
As with other grazers, they can be very selective, feeding on the leaves of one species and disregarding other species surrounding it. They eat a greater variety of plants during the dry season, as fewer plants are available. While they eat grass during the wet season, they have to switch to more abundant Reed (plant), reeds during the dry season. The capybara's jaw hinge is not perpendicular; hence, it chews food by grinding back-and-forth rather than side-to-side.Capybara. ''Hydrochaeris hydrochaeris''
San Francisco Zoo
Capybara are coprophagous as a means of obtaining bacterial gut flora to help digestion, digest the cellulose in the grass that forms their normal diet, and to extract the maximum protein and vitamins from their food. They may also regurgitate food to masticate again, similar to cud-chewing by a cow. As with other rodents, the front teeth of capybara grow continually to compensate for the constant wear from eating grasses;Bristol Zoo Gardens (UK) ''Capybara''
. Bristolzoo.org.uk. Retrieved on 2011-12-07.
their cheek teeth also grow continuously.


Pseudoruminant

The
hippopotamus The hippopotamus ( ; : hippopotamuses or hippopotami; ''Hippopotamus amphibius''), also called the hippo, common hippopotamus, or river hippopotamus, is a large semiaquatic mammal native to sub-Saharan Africa. It is one of only two extan ...
is a large, semi-aquatic, mammal inhabiting rivers, lakes and mangrove swamps. During the day, they remain cool by staying in the water or mud; reproduction and childbirth both occur in water. They emerge at dusk to graze on grasses. While hippopotamuses rest near each other in the water, grazing is a solitary activity. Their incisors can be as long as 40 cm and the Canine tooth, canines up to 50 cm; however, the canines and incisors are used for combat, and play no role in feeding. Hippos rely on their broad, horny lips to grasp and pull grasses which are then ground by the Molar (tooth), molars. The hippo is considered to be a
pseudoruminant Pseudoruminant is a classification of animals based on their digestive tract differing from the ruminants. Hippopotami and camels are ungulate mammals with a three-chambered stomach (ruminants have a four-chambered stomach) while equids (horses, as ...
; it has a complex three- or four-chambered stomach but does not "chew cud".


Non-grass grazing

Although grazing is typically associated with mammals feeding on grasslands, ecologists sometimes use the word in a broader sense, to include any organism that feeds on any other species without ending the life of the prey organism. Use of the term "grazing" varies further; for example, a marine biologist may describe herbivorous sea urchins that feed on kelp as ''grazers'', even when they kill the organism by cutting the plant at the base. Malacologists sometimes apply the word to aquatic snails that feed by consuming the microscopic film of algae, diatoms and detritus—a biofilm—that covers the substrate (biology), substrate and other surfaces underwater. In marine ecosystems, grazing by mesograzers such as some crustaceans maintains habitat structure by preventing algal overgrowth, especially in coral reefs.


Benefits


Environmental

Grazer urine and feces "recycle nitrogen, phosphorus, potassium and other plant nutrients and return them to the soil"."Benefits of Grazing Cattle on the Prairie". Native Habitat Organization. Retrieved 1 Dec 2008
Grazing can allow for the accumulation of organic matter which may help to combat soil erosion. This acts as nutrition for insects and organisms found within the soil. These organisms "aid in carbon sequestration and water filtration".


Biodiversity

When grass is grazed, dead litter grass is reduced which is advantageous for birds such as waterfowl. Grazing can increase biodiversity. Without grazing, many of the same grasses grow, for example Bromus, brome and Poa, bluegrass, consequently producing a monoculture.


References


External links

{{Authority control Ethology Herbivory Land use